its really a combination of two things... the 1+2, 3+4, 5 are treated the same from a defensive formula perspective. this means you could flip the 1 and 2 or the 3 and 4 and have no impact on your defensive prowess. but in the end, to some extent, all 5 players are averaged.
what we don't know is if that means they are averaged in equal weights or not. is the 1 as valuable as the 5 regarding shots from 20 feet? from 2 feet? its not really clear - my take is the guards get plugged into the guard formula (the 1+2 formula in the 2-3 zone) and probably return higher values, at 20 feet, than a center does in the formula for the 5. maybe the pg is great, worth 10 points, the sg is ok, worth 7.5, and the center is good - but only worth 5, because inherently he isn't as fast and/or the formula for the 5 at 20 feet just doesn't return as large of values.
but, its also possible that's not really true - and that a center is basically as important as a pg for any shot. in short, we know all 5 guys contribute to the defense on every shot. we don't know if they contribute equally. my best guess is there is some disparity, guards matter more for per shots, bigs for post shots, but that the disparity is moderate, not huge.